生活型

herb

形態特徵

• Evergreen creeping lycophytes, without erect or ascending stems.
• Stems 3-5 cm long, 2.5-3.5 mm wide (including microphylls), rarely longer than 10 cm, decumbent or creeping, undersides and uppersides differentiated, irregularly and dichotomously branched. Rhizophores present, short, originated from the upperside of stem at the branch site, 0.1-0.2 mm in diameter; axillary trophophylls present at branching points, lanceolate, ciliate along margin.
• Trophophylls conspicuously dimorphic, arranged in 4 ranks (2 dorsal & 2 ventral), margin more or less white-callous; ventral (lateral) trophophylls ovate, 1.2-2 mm long, usually less than 1 mm wide, attached by broad base, caudate at apex, loosely serrulate along margin but ciliate at acroscopically basal part; dorsal (median) trophophylls broadly lanceolate to ovate, 1.2-1.8 mm long, 0.6-0.8 mm wide, base rounded, apex caudate or long-tail, margin loosely serrulate, conspicuously white-callous. Strobili usually solitary, sometimes twin on the forked branches, resupinate, 0.4-1.5 cm long, megasporangia usually formed at basal sporophylls and microsporangia formed at upper ones; dorsal sporophylls folded, 2-2.5 mm long, lanceolate by one side, ciliate along margin of basal part, acuminate or tailed at apex, rounded at base; ventral sporophylls shovel-like, ovate or broadly ovate, keeled, ca. 1.6-2 mm long, 1-1.2 mm wide, ciliate along margin, acuminate or tailed at apex, rounded at base.
• Megaspore (Figs. 2E & F) anisopolar, radiosymmetrical, trilete, yellow, usually 4 in a megasporangium, equatorial diameter 280-300 μm; perispore rugulate or rugulo-reticulate, with well-defined irregular wrinkles. Microspore anisopolar, radiosymmetrical, trilete, reddish orange, numerous in a microsporangium, equatorial diameter 35-37 μm; perispore verrucate, with coral-like structures on the surface.

分布

• 台灣 : it grows on the moist rocky slope with some shade, and 4 populations were found in eastern Taiwan.
• 世界 : from Kyushu (Southern part of Kagoshima-ken) to Ryukyu Archipelago of Japan.

引證標本

Tatung County: Changpin Township, Pahsientung, 30-50 m alt., on the moist rock. P.-F. Lu 1311 (TAIF), C.-F. Chen s.n., Dec. 12, 2001 (TAIF), H.-M. Chang 6341 (TNU), Mt. Hsinkanshan, ca. 750 m alt., H.-M. Chang 6917 (TAIE, TAIF), Changyun, M.-Y. Sheng 4570 (TAIE); Lanyu, Yehyu, 10 m alt., on the laying stones, P.-F. Lu 8575 (TAIF), H.-M. Chang 6633 (TAIE).

參考文獻

• Ho-Ming Chang, Jinn-Fen Chen, Pi-Fong Lu, Shann-Jye Moore, Jenn-Che Wang and Wen-Liang Chiou. 2009. Taiwania 54(1): 88-92.